We use teleseismic body wave tomography to reveal anomalous P wave velocity variations in the upper mantle beneath south-east Australia. Data are sourced from the WOMBAT transportable seismic array, the largest of its kind in the southern hemisphere, which enables horizontal resolution of approximately 50 km to be achieved over a large region that includes Victoria, New South Wales and southern South Australia. In order to account for long-wavelength structure that is lost due to the use of multiple teleseismic datasets from adjacent arrays with non-overlapping recording periods, the AuSREM mantle model is includedas prior information in the inversion. Furthermore, AuSREM crust and Moho structure is explicitly included in the initial model in order to account for the presence of shallow heterogeneity which is poorly constrained by the teleseismic dataset. The P wave velocity model obtained from the joint inversion of WOMBAT teleseismic data represents a vast new resource on the seismic structure of the upper mantle beneath south-east Australia. One of the most striking features of the model is the presence of a north-dipping low-velocity anomaly beneath the Newer Volcanics province, a Quaternary intraplate basaltic province in western Victoria. The anomaly appears to terminate at approximately 200 km depth and has a structure that is more suggestive of a source confined to the upper mantle rather than a deeply rooted mantle plume. Other features that can be observed include a high-velocity anomaly beneath the Curnamona province and a high-velocity salient beneath the New England Orogen. Of particular interest is an extensive highvelocity anomaly beneath the Lachlan Orogen, which coincides almost exactly with thesurface expression of the Hay-Booligal Zone in the south, and extends northwards beneath the Macquarie Arc. The higher velocities beneath the Hay-Booligal Zone are consistent with the idea that it may be floored by a fragment of Proterozoic continental lithosphere that was once part of the east Gondwana margin, while the higher velocities beneath the Macquarie Arc may be related to its origin as an intra-oceanic arc.